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@ARTICLE{Vlijm:120446,
author = {R. Vlijm$^*$ and S. H. Kim and P. L. De Zwart and Y. Dalal
and C. Dekker},
title = {{T}he supercoiling state of {DNA} determines the handedness
of both {H}3 and {CENP}-{A} nucleosomes.190},
journal = {Nanoscale},
volume = {9},
number = {5},
issn = {2040-3372},
address = {Cambridge},
publisher = {RSC Publ.},
reportid = {DKFZ-2017-00875},
pages = {1862 - 1870},
year = {2017},
abstract = {Nucleosomes form the unit structure of the genome in
eukaryotes, thereby constituting a fundamental tenet of
chromatin biology. In canonical nucleosomes, DNA wraps
around the histone octamer in a left-handed toroidal ramp.
Here, in single-molecule magnetic tweezers studies of
chaperone-assisted nucleosome assembly, we show that the
handedness of the DNA wrapping around the nucleosome core is
intrinsically ambidextrous, and depends on the pre-assembly
supercoiling state of the DNA, i.e., it is not uniquely
determined by the octameric histone core. Nucleosomes
assembled onto negatively supercoiled DNA are found to
exhibit a left-handed conformation, whereas assembly onto
positively supercoiled DNA results in right-handed
nucleosomes. This intrinsic flexibility to adopt both
chiralities is observed both for canonical H3 nucleosomes,
and for centromere-specific variant CENP-A nucleosomes.
These data support recent advances suggesting an intrinsic
adaptability of the nucleosome, and provide insights into
how nucleosomes might rapidly re-assemble after cellular
processes that generate positive supercoiling in vivo.},
cin = {E190},
ddc = {600},
cid = {I:(DE-He78)E190-20160331},
pnm = {315 - Imaging and radiooncology (POF3-315)},
pid = {G:(DE-HGF)POF3-315},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:28094382},
doi = {10.1039/C6NR06245H},
url = {https://inrepo02.dkfz.de/record/120446},
}