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@ARTICLE{Yoo:120638,
author = {Y. Yoo and J. H. Park and C. Weigel$^*$ and D. B.
Liesenfeld and D. Weichenhan$^*$ and C. Plass$^*$ and D.-G.
Seo and A. M. Lindroth and Y. J. Park},
title = {{TET}-mediated hydroxymethylcytosine at the {P}parγ locus
is required for initiation of adipogenic differentiation.},
journal = {International journal of obesity and related metabolic
disorders},
volume = {41},
number = {4},
issn = {1476-5497},
address = {London [u.a.]},
publisher = {Nature Publ. Group56782},
reportid = {DKFZ-2017-01066},
pages = {652 - 659},
year = {2017},
abstract = {Adipose tissue is one of the main organs regulating energy
homeostasis via energy storage as well as endocrine
function. The adipocyte cell number is largely determined by
adipogenesis. While the molecular mechanism of adipogenesis
has been extensively studied, its role in dynamic DNA
methylation plasticity remains unclear. Recently, it has
been shown that Tet methylcytosine dioxygenase (TET) is
catalytically capable of oxidizing DNA 5-methylcytosine
(5-mC) to 5-hydroxymethylcytosine (5-hmC) toward a complete
removal of the methylated cytosine. We investigate whether
expression of the Tet genes and production of
hydroxymethylcytosine are required for preadipocyte
differentiation.Murine 3T3-L1 preadipocytes were used to
evaluate the role of Tet1 and Tet2 genes during
adipogenesis. Changes in adipogenic ability and in
epigenetic status were analyzed, with and without
interfering Tet1 and Tet2 expression using small interfering
RNA (siRNA). The adipogenesis was evaluated by Oil-Red-O
staining and induced expression of adipogenic genes using
quantitative polymerase chain reaction (qPCR). Levels of
5-hmC and 5-mC were measured by MassARRAY,
immunoprecipitation and GC mass spectrometry at specific
loci as well as globally.Both Tet1 and Tet2 genes were
upregulated in a time-dependent manner, accompanied by
increased expression of hallmark adipogenic genes such as
Pparγ and Fabp4 (P<0.05). The TET upregulation led to
reduced DNA methylation and elevated hydroxymethylcytosine,
both globally and specifically at the Pparγ locus (P<0.05
and P<0.01, respectively). Knockdown of Tet1 and Tet2
blocked adipogenesis (P<0.01) by repression of Pparγ
expression (P<0.05). In particular, Tet2 knockdown repressed
conversion of 5-mC to 5-hmC at the Pparγ locus (P<0.01).
Moreover, vitamin C treatment enhanced adipogenesis
(P<0.05), while fumarate treatment inhibited it (P<0.01) by
modulating TET activities.TET proteins, particularly TET2,
were required for adipogenesis by modulating DNA methylation
at the Pparγ locus, subsequently by inducing Pparγ gene
expression.},
cin = {C010},
ddc = {610},
cid = {I:(DE-He78)C010-20160331},
pnm = {322 - Genetics and Pathophysiology (POF3-322)},
pid = {G:(DE-HGF)POF3-322},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:28100914},
doi = {10.1038/ijo.2017.8},
url = {https://inrepo02.dkfz.de/record/120638},
}