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@ARTICLE{Schott:128288,
author = {J. Schott$^*$ and S. Reitter$^*$ and J. Philipp$^*$ and K.
Haneke$^*$ and H. Schäfer and G. Stoecklin$^*$},
title = {{T}ranslational regulation of specific m{RNA}s controls
feedback inhibition and survival during macrophage
activation.},
journal = {PLoS Genetics},
volume = {10},
number = {6},
issn = {1553-7404},
address = {San Francisco, Calif.},
publisher = {Public Library of Science},
reportid = {DKFZ-2017-04305},
pages = {e1004368 -},
year = {2014},
abstract = {For a rapid induction and efficient resolution of the
inflammatory response, gene expression in cells of the
immune system is tightly regulated at the transcriptional
and post-transcriptional level. The control of mRNA
translation has emerged as an important determinant of
protein levels, yet its role in macrophage activation is not
well understood. We systematically analyzed the contribution
of translational regulation to the early phase of the
macrophage response by polysome fractionation from mouse
macrophages stimulated with lipopolysaccharide (LPS).
Individual mRNAs whose translation is specifically regulated
during macrophage activation were identified by microarray
analysis. Stimulation with LPS for 1 h caused translational
activation of many feedback inhibitors of the inflammatory
response including NF-κB inhibitors (Nfkbid, Nfkbiz, Nr4a1,
Ier3), a p38 MAPK antagonist (Dusp1) and
post-transcriptional suppressors of cytokine expression
(Zfp36 and Zc3h12a). Our analysis showed that their
translation is repressed in resting and de-repressed in
activated macrophages. Quantification of mRNA levels at a
high temporal resolution by RNASeq allowed us to define
groups with different expression patterns. Thereby, we were
able to distinguish mRNAs whose translation is actively
regulated from mRNAs whose polysomal shifts are due to
changes in mRNA levels. Active up-regulation of translation
was associated with a higher content in AU-rich elements
(AREs). For one example, Ier3 mRNA, we show that repression
in resting cells as well as de-repression after stimulation
depends on the ARE. Bone-marrow derived macrophages from
Ier3 knockout mice showed reduced survival upon activation,
indicating that IER3 induction protects macrophages from
LPS-induced cell death. Taken together, our analysis reveals
that translational control during macrophage activation is
important for cellular survival as well as the expression of
anti-inflammatory feedback inhibitors that promote the
resolution of inflammation.},
keywords = {Adaptor Proteins, Signal Transducing (NLM Chemicals) /
Cytokines (NLM Chemicals) / IEX-1 protein, mouse (NLM
Chemicals) / Immediate-Early Proteins (NLM Chemicals) /
Lipopolysaccharides (NLM Chemicals) / NF-kappa B (NLM
Chemicals) / Nfkbiz protein, mouse (NLM Chemicals) / Nr4a1
protein, mouse (NLM Chemicals) / Nuclear Proteins (NLM
Chemicals) / Nuclear Receptor Subfamily 4, Group A, Member 1
(NLM Chemicals) / RNA, Messenger (NLM Chemicals) /
Tristetraprolin (NLM Chemicals) / Zfp36 protein, mouse (NLM
Chemicals) / p38 Mitogen-Activated Protein Kinases (NLM
Chemicals) / Ribonucleases (NLM Chemicals) / Zc3h12a
protein, mouse (NLM Chemicals) / Dual Specificity
Phosphatase 1 (NLM Chemicals) / Dusp1 protein, mouse (NLM
Chemicals)},
cin = {A200},
ddc = {570},
cid = {I:(DE-He78)A200-20160331},
pnm = {311 - Signalling pathways, cell and tumor biology
(POF3-311)},
pid = {G:(DE-HGF)POF3-311},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:24945926},
pmc = {pmc:PMC4063670},
doi = {10.1371/journal.pgen.1004368},
url = {https://inrepo02.dkfz.de/record/128288},
}