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@ARTICLE{Sun:144733,
author = {H. Sun and S. Rana and Z. Wang and K. Zhao and M.
Schnölzer$^*$ and J. Provaznik and T. Hackert and Q. Lv and
M. Zöller},
title = {{T}he {P}ancreatic {C}ancer-{I}nitiating {C}ell {M}arker
{CD}44v6 {A}ffects {T}ranscription, {T}ranslation, and
{S}ignaling: {C}onsequences for {E}xosome {C}omposition and
{D}elivery.},
journal = {Journal of oncology},
volume = {2019},
issn = {1687-8469},
address = {New York, NY},
publisher = {Hindawi Publ. Corp.},
reportid = {DKFZ-2019-02165},
pages = {3516973},
year = {2019},
abstract = {Pancreatic cancer-initiating cells (PaCIC) express CD44v6
and Tspan8. A knockdown (kd) of these markers hinders the
metastatic capacity, which can be rescued, if the cells are
exposed to CIC-exosomes (TEX). Additional evidence that
CD44v6 regulates Tspan8 expression prompted us to explore
the impact of these PaCIC markers on nonmetastatic PaCa and
PaCIC-TEX. We performed proteome, miRNA, and mRNA deep
sequencing analyses on wild-type, CD44v6kd, and Tspan8kd
human PaCIC and TEX. Database comparative analyses were
controlled by qRT-PCR, Western blot, flow cytometry, and
confocal microscopy. Transcriptome analysis of CD44 versus
CD44v6 coimmunoprecipitating proteins in cells and TEX
revealed that Tspan8, several signal-transducing molecules
including RTK, EMT-related transcription factors, and
proteins engaged in mRNA processing selectively associate
with CD44v6 and that the membrane-attached CD44
intracytoplasmic tail supports Tspan8 and NOTCH
transcription. Deep sequencing uncovered a CD44v6
contribution to miRNA processing. Due to the association of
CD44v6 with Tspan8 in internalization prone
tetraspanin-enriched membrane domains (TEM) and the
engagement of Tspan8 in exosome biogenesis, most
CD44v6-dependent changes were transferred into TEX such that
the input of CD44v6 to TEX activities becomes largely waved
in both a CD44v6kd and a Tspan8kd. Few differences between
CD44v6kd- and Tspan8kd-TEX rely on CD44v6 being also
recovered in non-TEM derived TEX, highlighting distinct TEX
delivery from individual cells that jointly account for
TEX-promoted target modulation. This leads us to propose a
model in which CD44v6 strongly supports tumor progression by
cooperating with signaling molecules, altering transcription
of key molecules, and through its association with the mRNA
processing machinery. The association of CD44v6 with Tspan8,
which plays a crucial role in vesicle biogenesis, promotes
metastases by transferring CD44v6 activities into TEM and
TEM-independently derived TEX. Further investigations of the
lead position of CD44v6 in shifting metastasis-promoting
activities into CIC-TEX may offer a means of targeting
TEX-CD44v6 in therapeutic applications.},
cin = {B100},
ddc = {610},
cid = {I:(DE-He78)B100-20160331},
pnm = {312 - Functional and structural genomics (POF3-312)},
pid = {G:(DE-HGF)POF3-312},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:31485223},
pmc = {pmc:PMC6702834},
doi = {10.1155/2019/3516973},
url = {https://inrepo02.dkfz.de/record/144733},
}