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@ARTICLE{Zhu:276949,
author = {C. Zhu and Y. M. Soto-Feliciano and J. P. Morris and C.-H.
Huang and R. P. Koche and Y.-J. Ho and A. Banito and C.-W.
Chen and A. Shroff and S. Tian and G. Livshits and C.-C.
Chen and M. Fennell and S. A. Armstrong and C. D. Allis and
D. F. Tschaharganeh$^*$ and S. W. Lowe},
title = {{MLL}3 regulates the {CDKN}2{A} tumor suppressor locus in
liver cancer.},
journal = {eLife},
volume = {12},
issn = {2050-084X},
address = {Cambridge},
publisher = {eLife Sciences Publications},
reportid = {DKFZ-2023-01226},
pages = {e80854},
year = {2023},
abstract = {Mutations in genes encoding components of chromatin
modifying and remodeling complexes are among the most
frequently observed somatic events in human cancers. For
example, missense and nonsense mutations targeting the mixed
lineage leukemia family member 3 (MLL3, encoded by KMT2C)
histone methyltransferase occur in a range of solid tumors,
and heterozygous deletions encompassing KMT2C occur in a
subset of aggressive leukemias. Although MLL3 loss can
promote tumorigenesis in mice, the molecular targets and
biological processes by which MLL3 suppresses tumorigenesis
remain poorly characterized. Here, we combined genetic,
epigenomic, and animal modeling approaches to demonstrate
that one of the mechanisms by which MLL3 links chromatin
remodeling to tumor suppression is by co-activating the
Cdkn2a tumor suppressor locus. Disruption of Kmt2c
cooperates with Myc overexpression in the development of
murine hepatocellular carcinoma (HCC), in which MLL3 binding
to the Cdkn2a locus is blunted, resulting in reduced H3K4
methylation and low expression levels of the locus-encoded
tumor suppressors p16/Ink4a and p19/Arf. Conversely,
elevated KMT2C expression increases its binding to the
CDKN2A locus and co-activates gene transcription. Endogenous
Kmt2c restoration reverses these chromatin and
transcriptional effects and triggers Ink4a/Arf-dependent
apoptosis. Underscoring the human relevance of this
epistasis, we found that genomic alterations in KMT2C and
CDKN2A were associated with similar transcriptional profiles
in human HCC samples. These results collectively point to a
new mechanism for disrupting CDKN2A activity during cancer
development and, in doing so, link MLL3 to an established
tumor suppressor network.},
keywords = {Humans / Animals / Mice / Liver Neoplasms: genetics / Liver
Neoplasms: pathology / Tumor Suppressor Protein p14ARF:
genetics / Carcinoma, Hepatocellular: genetics / Carcinoma,
Hepatocellular: pathology / Cyclin-Dependent Kinase
Inhibitor p16: genetics / Cyclin-Dependent Kinase Inhibitor
p16: metabolism / Chromatin / Carcinogenesis / MLL3 (Other)
/ cancer (Other) / cancer biology (Other) / chromatin
(Other) / human (Other) / liver cancer (Other) / mouse
(Other) / tumor suppressor (Other) / Tumor Suppressor
Protein p14ARF (NLM Chemicals) / Cyclin-Dependent Kinase
Inhibitor p16 (NLM Chemicals) / Chromatin (NLM Chemicals) /
CDKN2A protein, human (NLM Chemicals)},
cin = {F190},
ddc = {600},
cid = {I:(DE-He78)F190-20160331},
pnm = {316 - Infektionen, Entzündung und Krebs (POF4-316)},
pid = {G:(DE-HGF)POF4-316},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:37261974},
pmc = {pmc:PMC10279454},
doi = {10.7554/eLife.80854},
url = {https://inrepo02.dkfz.de/record/276949},
}