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@ARTICLE{Lenz:277141,
author = {M. Lenz and A. Eichler and P. Kruse and C. Galanis and D.
Kleidonas and G. Andrieux and M. Börries$^*$ and P.
Jedlicka and U. C. Müller and T. Deller and A. Vlachos},
title = {{T}he amyloid precursor protein regulates synaptic
transmission at medial perforant path synapses.},
journal = {The journal of neuroscience},
volume = {43},
number = {29},
issn = {0270-6474},
address = {Washington, DC},
publisher = {Soc.},
reportid = {DKFZ-2023-01303},
pages = {5290-5304},
year = {2023},
note = {2023 Jul 19;43(29):5290-5304},
abstract = {The perforant path provides the primary cortical excitatory
input to the hippocampus. Due to its important role in
information processing and coding, entorhinal projections to
the dentate gyrus have been studied in considerable detail.
Nevertheless, synaptic transmission between individual
connected pairs of entorhinal stellate cells and dentate
granule cells remains to be characterized. Here, we have
used mouse organotypic entorhino-hippocampal tissue cultures
of either sex, in which the entorhino-dentate (EC-GC)
projection is present and EC-GC pairs can be studied using
whole-cell patch clamp recordings. By using cultures of
wildtype mice, the properties of EC-GC synapses formed by
afferents from the lateral and medial entorhinal cortex were
compared and differences in short-term plasticity were
identified. Since the perforant path is severely affected in
Alzheimer's disease, we used tissue cultures of
amyloid-precursor protein (APP)-deficient mice to examine
the role of APP at this synapse. APP deficiency altered
excitatory neurotransmission at medial perforant path
synapses, which was accompanied by transcriptomic and
ultrastructural changes. Moreover, presynaptic but not
postsynaptic APP deletion through the local injection of
Cre-expressing adeno-associated viruses in conditional
APPflox/flox tissue cultures increased the neurotransmission
efficacy at perforant path synapses. In summary, these data
suggest a physiological role for presynaptic APP at medial
perforant path synapses that may be adversely affected under
altered APP processing conditions.SIGNIFICANCE STATEMENTThe
hippocampus receives input from the entorhinal cortex via
the perforant path. These projections to hippocampal dentate
granule cells are of utmost importance for learning and
memory formation. Although there is detailed knowledge about
perforant path projections, the functional synaptic
properties at the level of individual connected pairs of
neurons are not well understood. In this study, we
investigated the role of the amyloid precursor protein (APP)
in mediating functional properties and transmission rules in
individually connected neurons using paired whole-cell
patch-clamp recordings and genetic tools in organotypic
tissue cultures. Our results show that presynaptic APP
expression limits excitatory neurotransmission via the
perforant path, which could be compromised in pathological
conditions such as Alzheimer's disease.},
cin = {FR01},
ddc = {610},
cid = {I:(DE-He78)FR01-20160331},
pnm = {899 - ohne Topic (POF4-899)},
pid = {G:(DE-HGF)POF4-899},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:37369586},
doi = {10.1523/JNEUROSCI.1824-22.2023},
url = {https://inrepo02.dkfz.de/record/277141},
}