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@ARTICLE{Galka:288728,
author = {D. Galka and T. T. Ali and A. Bast and M. Niederleithinger
and E. Gerhardt and R. Motosugi and E. Sakata and M.
Knop$^*$ and T. F. Outeiro and B. Popova and G. H. Braus},
title = {{I}nhibition of 26{S} proteasome activity by α-synuclein
is mediated by the proteasomal chaperone {R}pn14/{PAAF}1.},
journal = {Aging cell},
volume = {23},
number = {5},
issn = {1474-9718},
address = {Oxford [u.a.]},
publisher = {Wiley-Blackwell},
reportid = {DKFZ-2024-00437},
pages = {e14128},
year = {2024},
note = {2024 May;23(5):e14128 / DKFZ-ZMBH Alliance},
abstract = {Parkinson's disease (PD) is characterized by aggregation of
α-synuclein (α-syn) into protein inclusions in
degenerating brains. Increasing amounts of aggregated α-syn
species indicate significant perturbation of cellular
proteostasis. Altered proteostasis depends on α-syn protein
levels and the impact of α-syn on other components of the
proteostasis network. Budding yeast Saccharomyces cerevisiae
was used as eukaryotic reference organism to study the
consequences of α-syn expression on protein dynamics. To
address this, we investigated the impact of overexpression
of α-syn and S129A variant on the abundance and stability
of most yeast proteins using a genome-wide yeast library and
a tandem fluorescent protein timer (tFT) reporter as a
measure for protein stability. This revealed that the
stability of in total 377 cellular proteins was altered by
α-syn expression, and that the impact on protein stability
was significantly enhanced by phosphorylation at Ser129
(pS129). The proteasome assembly chaperone Rpn14 was
identified as one of the top candidates for increased
protein stability by expression of pS129 α-syn. Elevated
levels of Rpn14 enhanced the growth inhibition by α-syn and
the accumulation of ubiquitin conjugates in the cell. We
found that Rpn14 interacts physically with α-syn and
stabilizes pS129 α-syn. The expression of α-syn along with
elevated levels of Rpn14 or its human counterpart PAAF1
reduced the proteasome activity in yeast and in human cells,
supporting that pS129 α-syn negatively affects the 26S
proteasome through Rpn14. This comprehensive study into the
alternations of protein homeostasis highlights the critical
role of the Rpn14/PAAF1 in α-syn-mediated proteasome
dysfunction.},
keywords = {26S proteasome (Other) / Parkinson's disease (Other) /
posttranslational modifications (Other) / proteasomal
chaperone (Other) / protein homeostasis (Other) / tandem
fluorescent protein timer (Other) / yeast (Other) /
α-Synuclein (Other)},
cin = {A260},
ddc = {610},
cid = {I:(DE-He78)A260-20160331},
pnm = {311 - Zellbiologie und Tumorbiologie (POF4-311)},
pid = {G:(DE-HGF)POF4-311},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:38415292},
doi = {10.1111/acel.14128},
url = {https://inrepo02.dkfz.de/record/288728},
}