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@ARTICLE{Bodensohn:300744,
author = {U. S. Bodensohn and B. Dünschede and C. Kuhlmann and K.
Kumari and R. Ladig$^*$ and C. Grefen and E. Schleiff and D.
Fernandez and D. Schünemann},
title = {{GET}3{B} is involved in chloroplast biogenesis and
interacts with the thylakoidal {ALB}3 and {ALB}4
insertases.},
journal = {Plant cell reports},
volume = {44},
number = {5},
issn = {0721-7714},
address = {Heidelberg [u.a.]},
publisher = {Springer},
reportid = {DKFZ-2025-00904},
pages = {108},
year = {2025},
abstract = {Proteomic, functional physiological analyses of get3b
mutant plants highlight GET3B's role in chloroplast
function. Genetic and interaction analyses indicate get3b
and srp54 as mutual potentiators that might share terminal
insertases. Protein targeting and insertion into membranes
are essential for cellular organization and organelle
function. The Guided Entry of Tail-anchored (GET) pathway
facilitates the post-translational targeting and insertion
of tail-anchored (TA) membrane proteins. Arabidopsis
thaliana has four GET3 homologues, including AtGET3B and
AtGET3D localized to chloroplasts. These photosynthetic
organelles possess complex membrane systems, and the
mechanisms underlying their protein targeting and membrane
biogenesis are not fully understood. This study conducted a
comprehensive proteomic analysis of get3b mutant plastids,
which displayed significant alterations. Fluorometric based
complex assembly as well as CO2 assimilation analyses
confirmed that disruption of GET3B function displayed a
significant impact on photosystem II assembly as well as
carbon fixation, respectively, indicating a functional role
in chloroplast biogenesis. Additionally, genetic
interactions were found between GET3B and the two component
STIC system, which cooperates with the cpSRP pathway which
is involved in the co-translational sorting of thylakoid
proteins. Further, physical interactions were observed
between GET3B and the C-terminus of ALB3 and ALB4 in vitro
and the full length proteins in vivo, indicating a role of
GET3B in protein targeting and membrane integration within
chloroplasts. These findings enhance our understanding of
GET3B's involvement in stromal protein targeting and
thylakoidal biogenesis.},
keywords = {Arabidopsis Proteins: metabolism / Arabidopsis Proteins:
genetics / Arabidopsis: metabolism / Arabidopsis: genetics /
Chloroplasts: metabolism / Thylakoids: metabolism /
Photosystem II Protein Complex: metabolism / Mutation / ALB3
(Other) / ALB4 (Other) / GET3B (Other) / STIC1 (Other) /
STIC2 (Other) / cpSRP54 (Other) / Arabidopsis Proteins (NLM
Chemicals) / ALBINO 3 protein, Arabidopsis (NLM Chemicals) /
Photosystem II Protein Complex (NLM Chemicals)},
cin = {B230},
ddc = {580},
cid = {I:(DE-He78)B230-20160331},
pnm = {312 - Funktionelle und strukturelle Genomforschung
(POF4-312)},
pid = {G:(DE-HGF)POF4-312},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:40299103},
pmc = {pmc:PMC12040988},
doi = {10.1007/s00299-025-03500-2},
url = {https://inrepo02.dkfz.de/record/300744},
}