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@ARTICLE{Hermann:300828,
author = {J. Hermann$^*$ and T. Bortecen$^*$ and R. Kalis and A.
Kowar$^*$ and C. Pechincha$^*$ and V. Vogt and M.
Schneider$^*$ and D. Helm$^*$ and J. Krijgsveld$^*$ and F.
Loayza-Puch$^*$ and J. Zuber and W. Palm$^*$},
title = {m{TORC}1 cooperates with t{RNA} wobble modification to
sustain the protein synthesis machinery.},
journal = {Nature Communications},
volume = {16},
number = {1},
issn = {2041-1723},
address = {[London]},
publisher = {Springer Nature},
reportid = {DKFZ-2025-00942},
pages = {4201},
year = {2025},
note = {DKFZ-ZMBH Alliance / #EA:A330#LA:A330#},
abstract = {Synthesizing the cellular proteome is a demanding process
that is regulated by numerous signaling pathways and RNA
modifications. How precisely these mechanisms control the
protein synthesis machinery to generate specific proteome
subsets remains unclear. Here, through genome-wide CRISPR
screens we identify genes that enable mammalian cells to
adapt to inactivation of the kinase mechanistic target of
rapamycin complex 1 (mTORC1), the central driver of protein
synthesis. When mTORC1 is inactive, enzymes that modify
tRNAs at wobble uridines (U34-enzymes), Elongator and
Ctu1/2, become critically essential for cell growth in vitro
and in tumors. By integrating quantitative nascent
proteomics, steady-state proteomics and ribosome profiling,
we demonstrate that the loss of U34-enzymes particularly
impairs the synthesis of ribosomal proteins. However, when
mTORC1 is active, this biosynthetic defect only mildly
affects steady-state protein abundance. By contrast,
simultaneous suppression of mTORC1 and U34-enzymes depletes
cells of ribosomal proteins, globally inhibiting
translation. Thus, mTORC1 cooperates with tRNA U34-enzymes
to sustain the protein synthesis machinery and support the
high translational requirements of cell growth.},
keywords = {Mechanistic Target of Rapamycin Complex 1: metabolism /
Mechanistic Target of Rapamycin Complex 1: genetics / RNA,
Transfer: metabolism / RNA, Transfer: genetics / Protein
Biosynthesis / Humans / Ribosomes: metabolism / Ribosomal
Proteins: metabolism / Ribosomal Proteins: genetics / HEK293
Cells / Animals / Proteomics / Uridine: metabolism /
Mechanistic Target of Rapamycin Complex 1 (NLM Chemicals) /
RNA, Transfer (NLM Chemicals) / Ribosomal Proteins (NLM
Chemicals) / Uridine (NLM Chemicals)},
cin = {A330 / B230 / B250 / W120},
ddc = {500},
cid = {I:(DE-He78)A330-20160331 / I:(DE-He78)B230-20160331 /
I:(DE-He78)B250-20160331 / I:(DE-He78)W120-20160331},
pnm = {311 - Zellbiologie und Tumorbiologie (POF4-311)},
pid = {G:(DE-HGF)POF4-311},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:40328729},
pmc = {pmc:PMC12056009},
doi = {10.1038/s41467-025-59185-4},
url = {https://inrepo02.dkfz.de/record/300828},
}