TY - JOUR AU - Niehrs, Christof AU - Seidl, Carina AU - Lee, Hyeyoon TI - An 'R-spondin code' for multimodal signaling ON-OFF states. JO - Bioessays VL - 46 IS - 10 SN - 0265-9247 CY - New York, NY PB - Wiley-Liss M1 - DKFZ-2024-01723 SP - e2400144 PY - 2024 N1 - #EA:A050#LA:A050# / DKFZ-ZMBH Alliance / 2024 Oct;46(10):e2400144 AB - R-spondins (RSPOs) are a family of secreted proteins and stem cell growth factors that are potent co-activators of Wnt signaling. Recently, RSPO2 and RSPO3 were shown to be multifunctional, not only amplifying Wnt- but also binding BMP- and FGF receptors to downregulate signaling. The common mechanism underlying these diverse functions is that RSPO2 and RSPO3 act as 'endocytosers' that link transmembrane proteins to ZNRF3/RNF43 E3 ligases and trigger target internalization. Thus, RSPOs are natural protein targeting chimeras for cell surface proteins. Conducting data mining and cell surface binding assays we report additional candidate RSPO targets, including SMO, PTC1,2, LGI1, ROBO4, and PTPR(F/S). We propose that there is an 'R-spondin code' that imparts combinatorial signaling ON-OFF states of multiple growth factors. This code involves the modular RSPO domains, notably distinct motifs in the divergent RSPO-TSP1 domains to mediate target interaction and internalization. The RSPO code offers a novel framework for the understanding how diverse signaling pathways may be coordinately regulated in development and disease. KW - BMP (Other) KW - FGF (Other) KW - Hedgehog (Other) KW - Rāspondin (Other) KW - Wnt (Other) KW - axon guidance (Other) KW - phosphatases (Other) KW - receptor internalization (Other) LB - PUB:(DE-HGF)16 C6 - pmid:39180250 DO - DOI:10.1002/bies.202400144 UR - https://inrepo02.dkfz.de/record/292456 ER -